A cosmopolitan inversion facilitates seasonal adaptation in overwintering Drosophila

Abstract
Fluctuations in the strength and direction of natural selection through time are a ubiquitous feature of life on Earth. One evolutionary outcome of such fluctuations is adaptive tracking, wherein populations rapidly adapt from standing genetic variation. In certain circumstances, adaptive tracking can lead to the long-term maintenance of functional polymorphism despite allele frequency change due to selection. 

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https://pubmed.ncbi.nlm.nih.gov/38051996/
Joaquin C B Nunez, Benedict A Lenhart, Alyssa Bangerter, Connor S Murray, Giovanni R Mazzeo, Yang Yu, Taylor L Nystrom, Courtney Tern, Priscilla A Erickson, Alan O Bergland